What does the cryptic crossword clue "Follow the track when he has departed (5)" mean?

The definition is "Follow". The wordplay uses a container (one word inside another). T (abbreviation="the") + RAIL (synonym="track") = TRAIL; definition: "Follow" The answer is Trail.

Follow the track when he has departed (5)

Telegraph Prize Cryptic #3128 | 10 Across | 2021-10-03 flower

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